17.1 Introduction

17.2 Phylogeny and Classification

17.3 Morphology

17.3.1 Shell

17.3.2 Head, Arms, and Funnel

17.3.2.1 Embryological Evidence for the Derivation of Head Structures

17.3.3 Epidermis (Skin)

17.3.4 Head and Funnel Retractor Muscles

17.3.4.1 Muscular Attachments, Shell Growth and the Addition of Septa

17.3.5 The Fossil Record of Soft Tissues

17.3.6 Mantle Wall

17.3.7 Mantle Cavity

17.4 Biology and Ecology

17.4.1 Habits and Habitats

17.4.1.1 The Habits and Habitats of Extinct Ectocochleate Cephalopods

17.4.2 Locomotion and Buoyancy

17.4.2.1 Locomotion

17.4.2.2 Buoyancy

17.4.2.3 The Shell as a Buoyancy Device

17.4.2.4 Other Buoyancy Strategies

17.4.3 The Ink Sac and Inking

17.4.4 Colour Changes – Chromatophores and Other Special Skin Structures

17.4.4.1 Chromatophores

17.4.4.2 Structural Reflectors (Iridophores, Silvery Reflectors, and Leucophores)

17.4.5 Luminescent Organs

17.5 Digestive System

17.5.1 The Jaws

17.5.2 The Radula

17.5.3 Feeding

17.6 Renopericardial System

17.7 Courtship, Reproduction, Development, and Growth

17.7.1 Reproductive System

17.7.1.1 Sexual Characters and Sexual Dimorphism

17.7.2 Courtship and Mating

17.7.3 Life History

17.7.3.1 Spawn and Parental Care

17.7.3.2 Development

17.7.4 The Nervous System, Sense Organs, and Behaviour

17.7.5 The Endocrine System

17.7.6 Learning and Intelligence

17.7.7 Behaviour

17.7.8 Symbiotic Relationships

17.7.9 Predators

17.8 Economic Values

17.9 The Origin of Cephalopods

17.10 Diversity and Fossil History

17.10.1 The Ordovician Radiation and Beyond

17.11 The Major Groups of Cephalopoda

17.11.1 The Nautiliforms

17.11.1.1 Ellesmeroceratia and Plectronoceratia

17.11.1.2 Orthoceratia

17.11.1.3 Discosorida

17.11.1.4 Endoceratia

17.11.1.5 Actinoceratia

17.11.1.6 Nautilia

17.12 Bactritia and Ammonitia (= Ammonoidea)

17.12.1 Bactritia (= Bactritoidea)

17.12.2 Ammonitia (Ammonites)

17.12.2.1 The Embryonic Shell (Ammonitella)

17.12.2.2 Ammonite Sutural Patterns

17.12.2.3 Life Habits

17.12.2.4 Aptychus and Jaws

17.12.2.5 Phylogeny, Classification, Diversity, and Fossil History of Ammonites

17.12.2.6 The Major Groups of Ammonites

17.13 Coleoida

17.13.1 Phylogeny, Classification, Diversity, and Fossil History

17.13.1.1 Belemnitidia (The Belemnites)

17.13.1.2 Decabrachia

17.13.1.3 Squid – A Grade Not a Clade – The Myopsida, Oegopsida, Bathyteuthoidea (‘Teuthida’)

17.13.1.4 Octobrachia (= Octopodiformes)

17.13.1.5 Giant Squid and Octopuses